![]() ![]() Polyclonal antibodies are in fact mixtures of antibodies with different affinities. Limitations of the measurement of the equilibrium constant At that point, the overall strength of the binding depends on the goodness of fit between the two surfaces (van der Waals forces are very short-range and decay with the seventh power of the interatomic distance 3) and their total contact area. At this distance, van der Waals forces prevail, but ionic groups still play a role. These attractive forces locally overcome the hydration energies of the two molecules, water molecules are expelled and the epitope and the paratope approach each other more closely. As presently understood, events at a molecular level occur as follows 2: when the epitope and the paratope casually come to a distance of several nanometres, they are attracted by long-range forces, such as ionic and hydrophobic bonds. Specific binding must overcome an overall repulsion between the two molecules. The specific binding between the antigenic determinant on the red cell (epitope) and the antigen-combining site on the immunoglobulin molecule (paratope) involves very small portions of the molecules 2, comprising just a few amino acids and a surface area between 0.4 and 8 nm 2. Weak interactions involved in epitope-paratope binding For a stable binding, several weak bonds must be present contemporaneously and this requires steric complementarity between the molecules 1. Therefore, even the strongest single weak interaction has an ephemeral life of a fraction of a second at physiological temperatures. Moreover, the kinetic energy is not distributed uniformly. At 37 ☌, water molecules have an average kinetic energies higher than the weakest bonds. These molecules are called “non-polar” because their atoms do not form electric dipoles. Actually, “hydrophobic” bonds are simply the result of the exclusion from water molecules of other molecules that cannot participate in good hydrogen bonds. Hydrogen bonds are very important in aqueous solutions because water easily forms strong hydrogen bonds. Hydrogen or ion-dipole bonds require oppositely charged atoms. Van der Waals forces are the weakest, but they are able to attract all kinds of molecules. A list of types of weak bonds is shown in table I, together with their strength (energy). According to quantum mechanics, all chemical bonds are based on electrostatic forces. ![]()
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